↑ Roberts, A. 1929. New forms of African mammals. Annals of the Transvaal Museum, 13: 91.
↑ Roberts, A. 1945. Descriptions of some new subspecies of mammals. Annals of the Transvaal Museum, 21: 65.
↑ Haas, S.K.; Hayssen, V.; Krausman, P.R.. Mammalian species - Panthera leo. 2005 Mammalian Species (762): 1-11. pdf
↑ Barnett, Ross; Nobuyuki Yamaguchi, Ian Barnes and Alan Cooper (August 2006). "Lost populations and preserving genetic diversity in the lion Panthera leo: Implications for its ex situ conservation". Conservation Genetics 7 (4): 507–14
↑ The Kruger Nationalpark Map. Honeyguide Publications CC. South Africa 2004.
↑ Barnett, R.; Yamaguchi, N.; Barnes, I.; and Cooper, A. (2006): The origin, current diversity and future conservation of the modern lion (Panthera leo). Proc. R. Soc. B (2006) 273, 2119–2125 doi:10.1098/rspb.2006.3555
